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Research papers

Changes in Rainfall Partitioning and Canopy Interception Modeling after Pro‐

gressive Thinning in Two Shrub Plantations in the Semiarid Loess Plateau in

China

Xiaotao Niu, Jun Fan, Mengge Du, Zijun Dai, Ruihua Luo, Hongyou Yuan,

Shougang Zhang

PII: S0022-1694(23)00241-X

DOI: https://doi.org/10.1016/j.jhydrol.2023.129299

Reference: HYDROL 129299

To appear in: Journal of Hydrology

Received Date: 23 May 2022

Revised Date: 25 December 2022

Accepted Date: 15 February 2023

Please cite this article as: Niu, X., Fan, J., Du, M., Dai, Z., Luo, R., Yuan, H., Zhang, S., Changes in Rainfall

Partitioning and Canopy Interception Modeling after Progressive Thinning in Two Shrub Plantations in the

Semiarid Loess Plateau in China, Journal of Hydrology (2023), doi: https://doi.org/10.1016/j.jhydrol.

2023.129299

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? 2023 Published by Elsevier B.V.

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Changes in Rainfall Partitioning and Canopy Interception Modeling after Progressive Thinning

in Two Shrub Plantations in the Semiarid Loess Plateau in China

Xiaotao Niu Conceptualization Investigation Methodology Software Visualization

Writing - original draft Writing - review & editinga,b,c,d, Jun Fan Conceptualization Data

curation Formal analysis Funding acquisition Methodology Project administration

Supervision Validation Writing - original draft Writing - review & editinga,b,d,,

fanjun@ms.iswc.ac.cn, Mengge Du Formal analysis Writing - review & editingd, Zijun Dai

Methodology Writing - review & editingd, Ruihua Luo Validation Investigationd,

Hongyou Yuan Investigationd, Shougang Zhang Investigationd

aThe Research Center of Soil and Water Conservation and Ecological Environment, Chinese Academy

of Sciences and Ministry of Education, Yangling, Shaanxi 712100, China

bInstitute of Soil and Water Conservation, Chinese Academy of Sciences and Ministry of Water

Resources, Yangling, Shaanxi 712100, China

cUniversity of Chinese Academy of Sciences, Beijing 100049, China

dState Key Laboratory of Soil Erosion and Dryland Farming on the Loess Plateau, Northwest A&F

University, Yangling, Shaanxi 712100, China

Corresponding author.

Highlights

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A five-year experiment with two thinning intensities in two dense shrub plantations.

Effects of thinning on rainfall partitioning and canopy parameters were quantified.

The observed drought year did not have significant impact in the HT plots.

Revised Gash model performed an underestimated interception in two shrubs (RE< 20%).

Gash model performed better in thinning plots than control plots.

Abstract

Understanding the interaction between the hydrological cycle and vegetation management strategies,

such as thinning, is essential to improve watershed management and support ecological services.

However, it remains unclear how thinning affects the key components of hydrological cycle of dense

shrublands in the semiarid regions. Thus, the purpose of this study was to analyze and evaluate the

effect of thinning on rainfall partitioning and interception simulations in dense shrublands. In a 5-year

field experiment, we compared rainfall partitioning in two re-vegetated shrublands (Caragana

korshinskii and Salix psammophila) in the Chinese Loess Plateau under two thinning intensities

(moderate thinning [MT] and heavy thinning [HT] refer to the removal of 25% and 50% of the

branches, respectively) or no thinning (NT) (control). We modeled canopy interception losses (I) in the

two thinning treatments and control treatment using the revised Gash model. The results showed that

under MT and HT, the throughfall (TF) rate increased by about 12% and 20%, respectively, compared

to NT. The stemflow (SF) and observed I rates decreased by about 26% and 33%, respectively, under

MT, and the corresponding values for HT were about 50% and 52%, respectively. The observed I rate

decreased proportional to the percentage of biomass removed from the C. korshinskii and S.

psammophila plots. The results also revealed a significant linear correlation between the plant area

index (PAI) value and the canopy water balance of the two shrub plantations (R2>0.83, P<0.05). The

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performance of the revised Gash model (i.e., relative error [RE] < 20%) was satisfactory according to

the Nash?Sutcliffe model efficiency (NSE) coefficient (0.34-0.71). Based on the RE values, the

performance of the revised Gash model was better when applied to the plots subjected to MT and HT

(RE≤3%) than NT (RE=9.3%) for S. psammophila. Changes in the canopy storage capacity and canopy

evaporation rate strongly affected changes in simulated interception loss. The model can facilitate

water management in semiarid shrub plantations by accurately simulating the effect of thinning on

interception loss.

Keywords: Rainfall partitioning, Interception, Revised Gash model, Thinning, Shrubland

1. Introduction

Vegetation restoration based on unreasonable density is a fact that currently exists in many arid and

semiarid areas (Cao et al., 2011; Farley et al., 2005; Feng et al., 2016; Molina & del Campo, 2012).

Highly dense vegetation consumes soil moisture and increases interception of limited precipitation in

arid and semiarid regions (Chen et al., 2010; Christina et al., 2017; Ge et al., 2022; Robinson et al.,

2006; Shao et al., 2018). As a result, insufficient water supply to supplement evapotranspiration, which

ultimately aggravates soil drought conditions and leads to the formation of a dry layer in the soil. The

conditions as mentioned above make the vegetation unsustainable, with vegetation decline (partial

death) or stunting, where trees remain small for decades (Jia et al., 2019; Navarro-Cerrillo et al., 2019;

Wang et al., 2010; Zhang et al., 2020). Under these conditions, the water conservation function of the

vegetation cover is extremely limited. Therefore, studying changes in key components of the

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hydrological cycle after reducing vegetation density in water-limited regions is necessary to improve

watershed management and support ecological services.

In arid and semi-arid areas of the Loess Plateau, like the northern areas of the Qinghai-Tibetian Plateau

and the Greater Hinggan Mountains in China, The vegetation in these areas is dominated by shrubs

(Zhang et al., 2022). Due to the nature conservation policy of the local government, these shrubs have

seldom been managed since their established, resulting in unmanaged dense shrub canopies that

intercept more rainfall and consume more water. Drought-induced branch dieback has recently been a

frequent phenomenon in these plantations (Ma et al., 2020). This suggests that the current structural

characteristics of these shrub regions may not be conducive to alleviating regional drought, and these

areas need more rainfall to preserve and supplement local water resources. It is necessary to allow as

much rainfall as possible to reach the surface and be stored in the soil, rather than being lost to

evaporation through interception. As a result, shrub plantations could be reduced in density at a large

spatial scale to increase resilience in the face of severe drought stress and future climate change.

Thinning is a common vegetation management practice aimed at reducing the canopy density and

improving the quality of the remaining vegetation (Gebhardt et al., 2014; Grunicke et al., 2020;

Momiyama et al., 2021; Sun et al., 2015). In recent decades, experimental evidence from di?erent

forest ecosystems has demonstrated that thinning can promote the vitality of residual trees and reduce

long-term stress due to competition for water, nutrients, and light. Thinning can also increase the

resilience and resistance of forest trees to severe drought stress and thus may be an effective approach

to climate adaptation (Gebhardt et al., 2014; Grunicke et al., 2020; Navarro-Cerrillo et al., 2019; Wang

et al., 2019). In addition, thinning affects hydrological processes, including evaporation, transpiration,

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rainfall partitioning, soil water content, and surface ?ow, in vegetated ecosystems (Gebhardt et al.,

2014; del Campo et al., 2019; Sun et al., 2014). Canopy interception, which accounts for 5?40% of

total rainfall, is an important part of the shrub water cycle (Yue et al., 2021). It determines the amount

of net rainfall reaching the soil and is a key component in the hydrological cycle. However, few studies

have explored practical strategies for managing highly dense shrublands, with most focusing on forests

(Grunicke et al., 2020; Sun et al., 2015).

In recent decades, many observational studies have focused on canopy interception of shrubs in arid

and semiarid areas at the individual plant scale (An et al., 2022; Domingo et al., 1998; Jian et al., 2018;

Li et al., 2008; Tonello et al., 2021; Yang et al., 2019a; Yuan et al., 2017; Zhang et al., 2015).

Extrapolating the results of these studies to heterogeneous landscapes is a challenge. In this context,

interception needs to be quantified at the stand or plot scale rather than at the individual plant scale due

to plot-level measurements of interception have the advantage that the results can be scaled up, for

example, using traditional cover methods (Snyder et al., 2021). Furthermore, plot-level experimental

results are generally applicable to a watershed, and they cannot be confidently applied where

conditions are markedly different, particularly regarding rainfall regimes and vegetation types. While

interception models allow the extrapolation of measurements in space and time and can be used to

predict the effects of climate and land cover change on water resources (Magliano et al., 2022).

Therefore, model simulations continue to be the main focus of vegetation canopy interception research.

Widely used models for estimating interception losses are the original Gash model and the revised

version of this model (Muzylo et al., 2009). The original Gash model, a simpli?ed version of the Rutter

model (Gash., 1979; Rutter et al., 1975). The Gash model represents rainfall input as a series of discrete

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storms separated by intervals suf?ciently long for the canopy and stems to dry completely—this

assumption is possible because of the rapid drying of forest canopies. Each storm is then divided into

three subsequent phases—canopy wetting-up, saturation, and drying. This separation emphasizes the

relative importance of the climate against plant structure. Gash et al. (1995) proposed the sparse

versions of Gash model to adjust the original model formulations to forest stands with signi?cant open

spaces between the tree canopies. A crucial change is that in the sparse versions, the evaporation rate

from wetted surfaces is no longer calculated for the entire plot area but only for the area covered by the

canopy. This change overcame a poor boundary condition in the original models whereby the modeled

canopy failed to wet up beyond a certain degree of sparseness. As might be expected, the Gash sparse

model gives better results than the original version in terms of modelling error. More than 76% of the

model applications resulted in errors below 10%, with an important contribution of model

performances with errors under 5% (51% of the applications). In contrast, the Gash original model the

figure was 27%. The better performance of the Gash sparse model may be due to the conceptual

changes introduced in this version but it may also be caused by many of the applications not being duly

validated. It should also be mentioned that the original version has mainly been applied to temperate

climates, whereas the sparse version has been applied mainly to tropical and semiarid climates, also

with good results (Deng et al., 2022; Herbst et al., 2006; Junqueira Junior et al., 2019; Limousin et al.,

2008; Ma et al., 2019; 2020). However, this model has rarely been applied to shrubs compared to

forests in semiarid areas. The lack of research in this area can be attributed to the fact that evaporation

from wet shrubland canopies, unlike wet forest canopies, is not a net water loss (David et al., 2005) and

because of the dif?cult techniques of water ?ow measurement (Dunkerley, 2000). Domingo et al. (1998)

showed that interception models could be successfully applied to shrubs, despite their structural

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differences from forests. Recently, several studies have measured rainfall partitioning after changes in

forest structure and derived interception parameters for model applications (Grunicke et al., 2020; Ma

et al., 2020; Sun et al., 2015). Most of these studies have been based on measurement data only over

several months, one growing season, or one year. Hence, the model parameters are derived from very

short datasets. Few long-term studies (i.e., > 1 year) have focused on shrub structural changes. Few

long-term studies (i.e., > 1 year) have focused on the effect of structural changes in the shrubland

canopy over time on interception loss.

The aims of this study were to (1) quantify the effects of vegetation changes on rainfall partitioning in

control and thinned plots of two shrub species (Caragana korshinskii and Salix psammophila) and (2)

clarify changes in interception processes using the revised Gash model based on the determination of

interception parameters. With these aims in mind, we conducted a 5-year experiment in which we

measured rainfall partitioning in two re-vegetated shrublands (C. korshinskii and S. psammophila) in

the Chinese Loess Plateau subjected to two thinning intensities (moderate thinning [MT] and heavy

thinning [HT]) and no thinning (NT). The revised Gash model modeled canopy interception losses in

the control and thinned treatments. This study’ results can help to predict and manage ecohydrological

change in water-limited ecosystems in the context of shifting shrub cover and climate conditions.

2. Methods

2.1 Study area

This study was conducted in the Liudaogou watershed (38.78°N, 110.35°E; 1,200 m altitude; 6.89 km2)

of Shenmu County, Shaanxi Province, China (Figure. 1). The watershed is located in an area of the

8

Loess Plateau known as a “water-wind erosion crisscross region.” The climate in the region is

mid-temperate semiarid. The average annual precipitation between 2003 and 2021 was 457 mm, with

approximately 83% of precipitation occurring between June and October. The potential evaporation is

1,200 mm y-1, and the average annual temperature is 9.6° C. January and July are the coldest and

warmest, with average monthly temperatures of -6.4 and 23.5° C, respectively. Aeolian sandy soil and

loess are typical soil types in this catchment. Two shrub species, C. korshinskii and S. psammophila,

are widely planted for ecological restoration in the catchment. Rainfall and high-temperature seasons

are synchronized, with plant growth in early April and plant senescence in late October.

The present experiment commenced in 2017, and monitoring continued until 2021. Two plantations of

C. korshinskii (30 × 10 m) and S. psammophila (15 × 6 m) on the level ground were selected. Each plot

was further subdivided into three equal plots of 10 × 10 m and 5 × 6 m (Figure. 1). C. korshinskii and S.

psammophila were planted in 2013, and the average plant heights at the start of the study were 165 and

290 cm, respectively. Both C. korshinskii and S. psammophila have an inverted-cone canopy and are

multi-stemmed shrubs with no trunk or branches extending obliquely from the base.

In June 2019, for C. korshinskii, one plot was subjected to MT, one was subjected to HT, and one was

left (NT) as a control. Likewise, for S. psammophila, one plot was subjected to MT, one was subjected

to HT, and one (NT) was left as a control (Figure. 1). In the MT plots, 25% of the branches were

removed, whereas 50% of the branches were removed in the HT plots. All thinning operations were

conducted by local villagers using branch shears to minimize soil disturbance on the plots. All twigs

and branches from the thinned shrubs were removed from the plots. The number of branches in each

plot post-treatment is shown in Figure 2. Despite the significant difference in the number of

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branches/base diameters in the three plots under the three various densities for two shrubs, the

frequency distribution was not significantly different.

2.2 Rainfall, throughfall, and stem?ow measurements

Gross precipitation (Pg, mm) was measured using an Onset? (Onset Computer Corp., Bourne, MA,

U.S.A.) RG3-M tipping bucket rain gauge (0.2 mm per tip) located 3 m outside the plots in the study

area. As automatic rain gauges generally underestimate rainfall, three manual rain gauges (20 cm

diameter) were placed around it for calibration and were read immediately after each rainfall event. In

this way, rainfall characteristics, such as the rainfall duration (h) and average rainfall intensity (mm h-1),

were calculated. A rain event was defined as a period with more than 0.2 mm of total Pg, separated by

at least 6 h without rain.

Throughfall (TF, mm) was measured using 13 rain gauges (20 cm diameter) at each study plot. To

overcome spatial variability in TF, the rain gauges at each plot were randomly placed every year. The

average TF was computed from all functioning rain gauges. To reduce evaporative loss from the rain

gauges, TF was measured within 2 h after rainfall ended during the daytime to reduce evaporative loss

from the rain gauges. If a rainfall event ended at night, TF was measured early the next morning.

Stem?ow (SF, mm) was measured on 12 representative branches at each study plot, with three

branches from branch basal diameter (BD) categories of < 5, 5?10, 10?15, and 15?18 mm. SF was

collected using aluminum foil collars. Each collar was fitted around the entire branch circumference

and near the base of the branch and sealed with neutral silicone caulk (Figure. 1). A 0.8 cm diameter

PVC hose was used to guide the SF into a container fitted with a lid. The collars and hoses were

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checked periodically for leaks and blockages, respectively. After obtaining these measurements, SF

was returned to the branch base to alleviate unnecessary drought stress on the sample branches. The SF

depth after each rainfall event was calculated in each shrub plot as follows:

(1)1=0

niVSF??

(2)dVPA

where SFv is the SF volume (L) of a plot after a rainfall event, Vi is the SF volume (ml) collected from

individual stems in a plot, n is the total number of stems in a plot, SFd is the SF depth (mm), and PA is

the plot area (m2).

The observed interception loss (I, mm) was calculated using Equation (3):

(3)g()IPTFS???

One-way ANOVA (post hoc Tukey''s tests) was used to recognize the significant differences (p < 0.05)

in rainfall partitioning variables across treatments in the same year and the same treatments in different

years. Statistical analysis was performed in SPSS (IBM SPSS Statistics, version 25.0).

2.3 Revised Gash analytical model

2.3.1 Description of the model

In the revised Gash analytical model, Pg is expressed as a series of discrete rainfall events in which the

canopy has sufficient time to dry before another rainfall event begins (Gash et al., 1995). Each rainfall

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event includes three phases: wetting, saturation, and cessation. The total canopy interception was

obtained by summing the interception losses of the canopy during each phase of a rainfall event.

Rainfall events were analyzed separately from a threshold value of Pg’ to determine whether the canopy

was saturated. To calculate interception loss, the revised analytical model incorporates two types of

data: canopy and climatic parameters. The canopy parameters are the canopy storage capacity (S),

canopy cover (c), free TF coe?cient (p) (calculated using Equation (4)), trunk storage capacity (St),

and rainfall fraction diverted into SF (pt). The climatic parameters are Pg, mean evaporation rate ( , E

mm h?1), and mean rainfall intensity ( , mm h?1) during each rainfall event. The amounts of rainwater R

required to saturate the canopy and trunk fully were calculated using Equations (5) and (6),

respectively:

(4)1pc??

(5)??g''ln/ccRPSE

(6)''/ttp?

where Pg’ and Pt’ are the amounts of rainwater required to saturate the canopy and trunk fully,

respectively; (mm h?1) is the mean evaporation rate per unit area of canopy cover; Sc (mm) is the cE

canopy storage capacity per unit area of cover.

The revised analytical model divides simulated interception loss into ?ve components, and the equation

used to calculate each component is shown in Table 1.

Table 1. Components of simulated interception loss in the revised Gash model.

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Components of simulated interception loss Formula

1 For m small storms insuf?cient to saturate the canopy (Pg < Pg’) ,1gimcP??

For n storms (Pg > Pg’) sufficient to saturate the canopy

2 Wetting the canopy ('')gcnS?

3 Wet canopy evaporation during storm events ??,''1giEPiR?

4 Evaporation after storm cessation cnS

5 Evaporation from trunks for q storms that saturate trunks (Pg > Pt’) ,1ttgiqqP????

2.3.2 Estimation of the model parameters

The S was obtained as a linear relationship between Pg and the sum of TF and SF, where the S was the

value of Pg when the sum of TF and SF was zero (Wallace & McJannet, 2008). Canopy cover was

estimated using hemispherical photographs taken by ?sheye webcams during the growing season, and

data were processed using CAN-EYE software (version 6.3) (Niu et al., 2021; Weiss & Baret, 2014).

The St and pt were estimated using the method of Gash and Morton (1978) as the negative intercept and

slope of the linear regression between Pg and SF. The was estimated using the method of Buttle E

and Farnsworth (2012) and calculated using Equation (7):

, (7) =EaR?

where “a” is the slope of the linear regression between Pg and observed I for Pg ≥ Pg’ and R

represents the mean rainfall intensity is less than 10 mm h-1.

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2.3.3 Model evaluation

The performance of the revised Gash analytical model was evaluated using relative error (RE, %), the

Nash?Sutcliffe model ef?ciency (NSE) metric (Nash & Sutcliffe, 1970), and the root mean square error

(RMSE). The RE (%) was the difference between the values estimated by the model and the measured

values. Muzylo et al. (2009) classified the performance of the interception rainfall model according to

the RE as follows: poor (RE > 30%), fair (10% < RE ≤ 30%), good (5% < RE ≤ 10%), very good (1%

< RE ≤ 5%), and extremely good (RE ≤ 1%). The NSE measures the performance of each interception

model as compared to the mean and can be used to assess the predictive power of the model. The

RMSE was used to quantify the agreement between the measured data and the model predictions.

These three metrics are more suitable used to evaluate hydrological models.

2.3.4 Sensitivity analysis

To explore the relative importance of the parameters in the revised Gash model, ?ve parameters (S, c,

Pt, St, and ) were subjected to a sensitivity analysis. In the analysis, the values of these /ER

parameters were increased or decreased by up to 50% of their original values. The simulated results

were then compared with the actual measurement data.

The methodology in this study is further presented in a flowchart in Figure 3.

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Figure 1. The study area in the Liudaogou catchment of Shenmu County, Shaanxi Province, China (a)

and C. korshinskii and S. psammophila plots subjected to no thinning (NT), moderate thinning (MT), or

heavy thinning (HT) by branch removal (b). Collection of gross rainfall (Pg) outside the plots (c) and

collection of throughfall (TF) (d) and stemflow (SF) (e).

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Figure 2. The number and frequency distributions of branch basal diameter (BD) in the shrub plots

under no thinning (NT), moderate thinning (MT), and heavy thinning (HT) in 2019.

Figure 3. Methodological flowchart.

3. Results

3.1 Rainfall characteristics

The daily precipitation amounts, intensities, and durations for the 2017?2021 are shown in Figure 4. In

total, 674.2 mm of precipitation occurred in 2017, 637.2 mm in 2018, 436.5 mm in 2019, 359.3 mm in

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2020, and 320.0 mm in 2021. According to the average annual precipitation (457.0 mm) between 2003

and 2021, 2017 and 2018 were classified as wet years, whereas 2019, 2020, and 2021 were classified as

normal, dry, and extremely dry years, respectively. Rainfall was unevenly distributed throughout the

year, with a mean of 88.0% occurring during the rainy season (May?October) in 2017?2021.

Individual rainfall events ranged from 0.2 to 92.5 mm, with an average of 7.6 mm. The rainfall

intensity ranged from 0.2 to 40.7 mm h-1, with an average of 4.2 mm h-1. The rainfall duration ranged

from 0.1 to 26.6 h, with an average of 3.1 h. As shown in Table 2, rainfall events less than 10 mm

account for about 75% of the rainfall events during 2017-2021, and about 90% have rainfall intensities

less than 10 mm h-1. The wet years are mainly characterized by a few more rainfall events greater than

40 mm compared to the dry years.

During the experimental periods (purple dashed lines, Figure. 4a) in the 5 years from 2017 to 2021, 116

rainfall events were recorded, with a total of 1,587.8 mm. Among them, 780.4 mm (49.1% of total Pg)

in the pre-thinning period (2017?2018) and 807.4 mm (50.9% of total Pg) in the post-thinning period

(2019?2021). The frequency distributions of event size and intensity during the pre-thinning and

post-thinning periods are shown in Figure 5. Generally, small rainfall events (low depths) were more

frequent and contributed to a lower total Pg percentage than large rainfall events (Figure. 5a, c).

Generally, the event frequency of low-intensity rainfall events (range: 0?5 mm h-1) was higher (> 50%

of total events) than the frequency of high-intensity rainfall events in the pre-thinning and post-thinning

periods (Figure. 5b, d). This suggested that lower-intensity rainfall events accounted for a higher

percentage of total Pg.

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Figure 4. Daily precipitation distribution for 2017?2021 (purple dashed lines = the experimental

periods).

18

Figure 5. Frequency distribution of rainfall events in di?erent ranges of rainfall amounts (a, c) and

rainfall intensities (b, d) during both the pre-thinning (2017?2018) and post-thinning (2019?2021)

periods in the study region.

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Table 2. Rainfall classification according to rainfall amount and intensity during observation years.

Rainfall classification Observation years n <2 2-5 5-10 10-20 20-40 >40

Amount (mm) 2017 86 34 (39.5%) 25 (29.1%) 8 (9.3%) 11 (12.8%) 3 (3.5%) 5 (5.8%)

2018 64 28 (43.8%) 6 (9.4%) 11 (17.2%) 9 (14.1%) 6 (9.3%) 4 (6.2%)

2019 61 25 (41.0%) 11 (18.0%) 13 (21.3%) 7 (11.5%) 3 (4.9%) 2 (3.3%)

2020 59 28 (47.4%) 13 (22.0%) 6 (10.2%) 7 (11.9%) 5 (8.5%) 0

2021 48 20 (41.7%) 8 (16.6%) 8 (16.6%) 9 (18.8%) 3 (6.3%) 0

2017-2021 318 135 (42.4%) 63 (19.8%) 46 (14.5%) 43 (13.5%) 20 (6.3%) 11 (3.5%)

Intensity (mm h-1) 2017 86 46 (53.5%) 21 (24.4%) 9 (10.4%) 6 (7.0%) 4 (4.7%) 0

2018 64 30 (46.9%) 23 (35.9%) 5 (7.8%) 4 (6.3%) 2 (3.1%) 0

2019 61 30 (49.2%) 20 (32.8%) 5 (8.2%) 3 (4.9%) 3 (4.9%) 0

2020 59 37 (62.7%) 12 (20.3%) 2 (3.4%) 3 (5.1%) 5 (8.5%) 0

2021 48 30 (62.5%) 11 (22.9%) 4 (8.3%) 1 (2.1%) 2 (4.2%) 0

2017-2021 318 173 (54.4%) 87 (27.4%) 24 (7.5%) 18 (5.7%) 16 (5.0%) 0

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3.2 Rainfall partitioning measurements

The changes in annual rainfall partitioning during the pre- and post-thinning periods are shown in

Figure 6 and Table 3. In 2017?2018, before thinning, there were no significant differences in the ratios

of TF, SF, and I to Pg in the C. korshinskii and S. psammophila plots subjected to NT, MT, and HT. In

2019?2021, after thinning, both thinned plots had a higher TF rate than the plots subjected to NT

(Figure. 6a, b). In both the C. korshinskii and S. psammophila plots, the TF rate increased by about 12%

(MT) and 20% (HT) compared to the plots subjected to NT. In contrast, the SF and I rates significantly

decreased with thinning intensity (Figure. 6c, d, e, f). In the C. korshinskii plots, the total SF decreased

by 21% (MT) and 49% (HT), and the total I decreased by 37% (MT) and 54% (HT) compared to the

plot subjected to NT. Corresponding values for S. psammophila were 31% (MT) and 50% (HT) for the

SF rate and 29% (MT) and 50% (HT) for the I rate. For both shrub species, although the ratio of I to Pg

was highest under the NT treatment in 2021, the plots in the thinned treatments had already reached

68.5% (MT) and 48.6% (HT) of this value for C. korshinskii, 73.0% (MT) and 67.2% (HT) for S.

psammophila. It is important to emphasize that in the plots subjected to HT, the effects of the

extremely dry year (2021) on total I cannot be determined.

The relationship between Pg and canopy water balance (TF, SF, and I) based on rainfall event data is

shown in Figure 7. The event-based canopy water balance amount increased linearly with Pg (P < 0.01).

The ratio of canopy water balance to Pg varied greatly throughout the study period and stabilized

gradually under rainfall events greater than 20 mm.

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Figure 6. Annual ratio of throughfall (TF), stemflow (SF), and canopy interception loss (I) to gross

rainfall (Pg) in the C. korshinskii and S. psammophila plots before (2017?2018) and after thinning

(2019?2021). In the figure, a capital letter indicates a signi?cant difference between the thinning

treatments (NT, MT, or HT), and a lower-case letter indicates a signi?cant difference between years

within the same treatment (p < 0.05).

22

Table 3. Observed rainfall amount, throughfall, stemflow, and interception loss during pre- (2017-2018) and post-thinning (2019-2021) periods in control and treated shrub

plots (2017 and 2018 were wet years, 2019, 2020, and 2021 were normal, dry, and extremely dry years, respectively).

Shrub species Plots Study period Rainfall amount (mm) Throughfall (mm) Stemflow (mm) Interception (mm)

C. korshinskii NT Pre-thinning 780.4 560.9 (71.9%) 77.1 (9.9%) 142.4 (18.2%)

Post-thinning 807.4 605.3 (75.0%) 80.3 (9.9%) 121.8 (15.1%)

MT Pre-thinning 780.4 553.7 (71.0%) 77.9 (9.9%) 148.8 (19.1%)

Post-thinning 807.4 667.6 (82.7%) 61.5 (7.6%) 78.3 (9.7%)

HT Pre-thinning 780.4 562.7 (72.1%) 75.7 (9.7%) 142.0 (18.2%)

Post-thinning 807.4 708.7 (87.8%) 41.9 (5.2%) 56.8 (7.0%)

S.psammophila NT Pre-thinning 780.4 553.8 (71.0%) 55.0 (7.0%) 171.6 (22.0%)

Post-thinning 807.4 601.2 (74.5%) 56.8 (7.0%) 149.4 (18.5%)

MT Pre-thinning 780.4 554.9 (71.1%) 55.2 (7.1%) 170.3 (21.8%)

Post-thinning 807.4 661.2 (81.9%) 39.9 (4.9%) 106.3 (13.2%)

HT Pre-thinning 780.4 554.1 (71.0%) 54.5 (7.0%) 171.8 (22.0%)

Post-thinning 807.4 696.7 (86.3%) 30.4 (3.7%) 80.3 (10.0%)

23

Figure 7. Relationship between gross rainfall (Pg) and canopy water balance using rainfall event data

from 2019 to 2021 in the C. korshinskii and S. psammophilas plots subjected to no thinning (NT),

moderate thinning (MT), or heavy thinning (HT). (a) Throughfall (TF) amount, (b) stemflow (SF)

amount, (c) canopy interception loss (I) amount, (d) ratio of TF to Pg, (e) ratio of SF to Pg, and (f) ratio

of I to Pg for C. korshinskii. (g) TF amount, (h) SF amount, (i) I amount, (j) ratio of TF to Pg, (k) ratio

of SF to Pg, and (l) ratio of I to Pg for S. psammophila. p < 0.05, p < 0.01

24

3.3 Model parameterization

Model input parameters may provide insights into the quality of the model’s performance and be useful

in the model’s result interpretation, so they should be given priority consideration. From the linear

regressions of Pg and rainfall partitioning components (Figure. 6), the study derived the required

parameter values in the revised Gash model applied to the C. korshinskii and S. psammophila plots

under NT, MT, and HT are shown in Tables 4 and 5. The estimated canopy parameters (S, PAI, c, Pt,

and St) and climatic parameter ( ) gradually decreased as thinning intensity increased. For example, E

the S of the C. korshinskii and S. psammophila plots decreased from 0.90 ± 0.05 to 0.40 ± 0.10 mm,

and the decreased from 0.30 ± 0.02 to 0.13 ± 0.03. Speci?cally, the S, Pt, and St values of C. E

korshinskii were slightly larger than those of S. psammophila in the plots under NT, MT, and HT.

Parameters for the observed periods agree with the development of vegetation structure (Table 4). The

thinning treatments reduced the measured parameters of vegetation structure, plant area index (PAI)

and c, to about 73% (MT) and 50% (HT) of the corresponding values of the control (NT) (Table 4). In

2017?2018 (two wet years), the relative PAI increase (rPAI-I, i.e., the change in the PAI divided by the

initial PAI) was within 10% of all plots. In 2019?2020 (a normal and dry year, respectively), the rPAI-I

declined to a lesser extent in the thinned plots (MT and HT) compared with the plots subjected to NT,

especially in the C. korshinskii plots. In 2021, under particularly dry conditions, the degree of decrease

in the rPAI-I of the plots subjected to HT was reduced by about half compared with that of the plots

under NT and MT. Although the PAI of the NT treatment was the highest in 2021, the plots on the

thinned treatments had already reached about 85% (MT) and 70% (HT) of this value for two shrubs.

Based on 5 years of measurement data, there was a simple linear relationship between the PAI values

25

and canopy water balance of the two shrub plantations (Figure. 8). According to the findings, the TF

rate decreased in accordance with an increase in the PAI in the two shrub plantations, whereas SF and I

rates increased in accordance with an increase in the PAI.

26

Table 4. The parameter values in the revised Gash analytical model applied to the C. korshinskii and S. psammophila plots subjected to no thinning (NT), moderate thinning

(MT), or heavy thinning (HT) during 2017?2021. Data obtained in the NT plots of the C. korshinskii and S. psammophila in 2017?2019 were used for calibration, and data

obtained in these plots in 2020?2021 were used for validation.

C. korshinskii S.psammophilaPlots Years

S (mm) PAI c p Pt St (mm) TF—E



R S (mm) PAI c p P

t St (mm) TF



E

NT 2017 0.82 2.33 0.85 0.15 0.100 0.121 0.24 2.30 0.78 1.64 0.71 0.29 0.078 0.098 0.35

2018 0.94 2.64 0.92 0.08 0.105 0.128 0.28 2.38 0.87 1.80 0.76 0.24 0.072 0.100 0.36

2019 0.85 2.52 0.90 0.10 0.108 0.119 0.26 2.43 0.85 1.77 0.73 0.27 0.076 0.106 0.33

2020 0.82 1.99 0.79 0.21 0.094 0.109 0.24 1.69 0.80 1.62 0.70 0.30 0.082 0.108 0.32

2021 0.68 1.47 0.65 0.35 0.074 0.087 0.22 2.02 0.58 1.16 0.60 0.40 0.059 0.077 0.29

MT 2017 0.86 2.35 0.83 0.17 0.103 0.112 0.23 2.30 0.76 1.65 0.70 0.30 0.078 0.094 0.34

2018 0.97 2.55 0.91 0.09 0.103 0.121 0.29 2.38 0.81 1.79 0.72 0.28 0.079 0.107 0.35

2019 0.65 1.74 0.65 0.35 0.080 0.099 0.17 2.43 0.60 1.27 0.53 0.47 0.051 0.075 0.29

2020 0.60 1.64 0.64 0.36 0.079 0.106 0.14 1.69 0.58 1.21 0.52 0.48 0.050 0.075 0.28

2021 0.48 1.27 0.55 0.45 0.058 0.076 0.14 2.02 0.40 0.86 0.45 0.55 0.043 0.065 0.25

HT 2017 0.86 2.34 0.82 0.18 0.099 0.111 0.25 2.30 0.75 1.67 0.70 0.30 0.075 0.090 0.36

2018 0.98 2.49 0.92 0.08 0.102 0.116 0.26 2.38 0.84 1.78 0.75 0.25 0.076 0.096 0.38

2019 0.42 1.14 0.47 0.53 0.060 0.086 0.13 2.43 0.38 0.87 0.38 0.62 0.042 0.071 0.22

2020 0.41 1.07 0.45 0.55 0.058 0.084 0.12 1.69 0.32 0.85 0.37 0.63 0.041 0.070 0.21

2021 0.40 0.93 0.44 0.56 0.048 0.072 0.11 2.02 0.30 0.76 0.35 0.65 0.037 0.063 0.19

27

Table 5. The calibration parameters in the revised Gash model for the C. korshinskii and S.

psammophila plots under the NT, MT, and HT treatments. The data were derived from June, July, and

September 2019?2021 datasets.

Shrubs Plots S (mm) c Pt St (mm) TF—E

C. korshinskii NT 0.75 0.74 0.090 0.105 0.28

MT 0.52 0.59 0.065 0.085 0.14

HT 0.40 0.44 0.050 0.075 0.10

S.psammophila NT 0.76 0.68 0.075 0.100 0.32

MT 0.46 0.48 0.046 0.070 0.22

HT 0.32 0.37 0.038 0.065 0.16

Figure 8. Relationship between plant area index (PAI) values and canopy water balance of the C.

korshinskii and S. psammophila plots for 2017?2021. The ratio of (a) TF to Pg, (b) SF to Pg, and (c) I to

Pg. p < 0.05

3.4. Rainfall interception modeling

The performance of the revised Gash model for the entire monitoring period (2017?2021) in the C.

korshinskii and S. psammophila plots subjected to NT is illustrated in Figure 9 and Table 6. Comparing

the simulated I with the measured values showed that the revised Gash model severely underestimated

I in wet years (2017 and 2018), especially in the presence of an individual rainfall event greater than 40

mm, such as that occurred in 2019, despite this classified as a normal year. In dry years (2020 and

28

2021), when there were many individual rainfall events of less than 10 mm, the revised Gash model

overestimated I. Applying the method proposed by Muzylo et al. (2009), the performance of the revised

Gash model in simulating I in both shrub plantations was classified as “fair” for both shrubs throughout

the observation period, “poor” in wet and extremely dry years, and “fair” in normal and dry years. But

in 2020 (dry year), the performance of the revised model in simulating I by S. psammophila was

classified as “very good.”

In the NT, MT, and HT plots of C. korshinskii and S. psammophila, individual rainfall events observed

during June, July, and September of 2019?2021 were used to calibrate the revised Gash model. The

calibrated models were then validated using measurements obtained in August and October 2019?2021.

The measured and simulated total I according to the revised Gash model are summarized in Table 7

and plotted in Figure 10. The I was underestimated to varying degrees in the calibration period, such as

in NT plots, with underestimation of 3.0% for C. korshinskii and 14.5% for S. psammophila. As the

degree of thinning increased, the degree of underestimation gradually increased. In contrast to the

results of the revised Gash model in the calibration period, in the validation period, the revised model

overestimated I, with the degree of overestimation decreasing in accordance with the thinning intensity.

Both root mean square error (RMSE) and NSE values gradually decreased with an increase in the

thinning intensity in both the calibration and validation periods. The performance of the revised Gash

model in all the plots varied from “fair” to “good” to “very good,” with a RE of < 20.0%. As shown in

Figure 10, the simulated values of individual rainfall events were slightly higher than the measured

values when the rainfall amounts were small. When the rainfall amounts were large, the simulated

values were lower than the measured values. Based on the correlation (R2) of the simulated and

observed values, the validation period was slightly better than the calibration period (Figure. 10).

29

Changes in interception components were compared in the six plots of the two shrub species (Figure.

11). The sum of evaporation during and after rainfall consistently accounted for the largest component

of estimated interception loss (about 90%). With an increase in the thinning intensity, evaporation

during rainfall gradually decreased in the C. korshinskii plots (from 51.6% to 42.9%) but increased in

the S. psammophila plots (from 55.1% to 57.1%). In contrast, evaporation after rainfall gradually

increased in the C. korshinskii plots (from 37.4% to 44.8%) but decreased in the S. psammophila plots

(from 35.4% to 30.6%).

To explore the relative importance of the parameters in the revised Gash model, a sensitivity analysis

of ?ve parameters (S, c, Pt, St, and ) was conducted (Figure. 12). and S were the most /ER/ER

sensitive parameters in the C. korshinskii and S. psammophila plots, followed by c, St, and Pt.

Figure 9. Accumulated total observed and simulated interception loss during the observation period

(2017?2019 data for calibration and 2020?2021 data for validation) in the C. korshinskii and S.

psammophila plots under the no thinning (NT) treatment according to the revised Gash model.

30

Table 6. Comparison of observed and simulated interception loss during the observation period

(2017?2021) in the C. korshinskii and S. psammophila plots under the no thinning (NT) treatment

according to the revised Gash analytical model.

Years Plots Observed I (mm) Simulated I (mm) RE (%) Classification

C. korshinskii NT 265.8 223.2 -16.0 Fair2017-2021

S. psammophila

NT

319.1 255.1 -20.0 Fair

C. korshinskii NT 70.4 38.6 -45.1 Poor2017

(calibration) S. psammophila

NT

69.6 44.0 -36.8 Poor

C. korshinskii NT 69.4 46.0 -33.7 Poor2018

(calibration) S. psammophila

NT

94.8 55.8 -41.1 Poor

C. korshinskii NT 59.4 43.2 -27.3 Fair2019

(calibration) S. psammophila

NT

66.5 49.2 -25.9 Fair

C. korshinskii NT 45.3 56.2 24.0 Fair2020

(validation) S. psammophila

NT

61.8 63.1 2.2 Very good

C. korshinskii NT 21.3 39.1 84.0 Poor2021

(validation) S. psammophila

NT

26.5 43.0 62.5 Poor

31

Figure 10. Observed interception loss versus simulated interception loss at the event-based scale

according to the revised Gash model in the calibration (June, July, and September 2019?2021) and

validation (August and October 2019?2021) periods in the C. korshinskii and S. psammophila plots

under no thinning (NT), moderate thinning (MT), or heavy thinning (HT).

32

Table 7. Comparison of observed and simulated interception loss according to the revised Gash analytical model in the calibration (June, July, and September 2019?2021)

and validation (August and October 2019?2021) periods in the C. korshinskii and S. psammophila plots under no thinning (NT), moderate thinning (MT), or heavy thinning

(HT).

Calibration period Validation period

C. korshinskii S.psammophila C. korshinskii S.psammophilaComponents of interception

NT MT HT NT MT HT NT MT HT NT MT HT

Simulated I (mm) 72.1 46.6 34.0 82.3 54.0 39.9 60.2 34.2 25.5 63.9 44.8 31.6

Observed I (mm) 74.3 50.4 36.8 96.2 66.8 49.7 51.7 29.5 22.1 58.4 43.5 31.1

Relative error (%) -3.0 -7.5 -7.6 -14.5 -19.2 -19.7 16.4 15.9 15.4 9.3 3.0 1.4

RMSE (mm) 1.34 0.45 0.23 1.35 1.19 0.87 1.68 0.25 0.13 0.81 0.69 0.43

NSE 0.44 0.36 0.34 0.48 0.39 0.38 0.49 0.47 0.46 0.71 0.70 0.67

Classification Very Good Good Good Fair Fair Fair Fair Fair Fair Good Very Good Very good



33

Figure 11. Interception components were estimated using the revised Gash model for 2019?2021 in

control (NT) and thinned (MT and HT) shrub plots.

34

Figure 12. Sensitivity analysis of the canopy parameters c, S, Pt, and St and climatic parameter /ER

in the revised Gash model under no thinning (NT), moderate thinning (MT), or heavy thinning (HT) in

the C. korshinskii and S. psammophila plots. Note: The change in c should be less than or equal to 1.

4. Discussion

4.1 Thinning effects on rainfall partitioning

The TF rate gradually increased, while the SF and observed I rates decreased with increasing thinning

intensity on an annual scale (Figure. 6). Although the gross amount of incident rainfall directly

determines the magnitudes of TF and SF (Figure. 7) and levels of saturation of canopy and stem

surfaces (Carlyle-Moses, 2004; Levia et al., 2010), many studies have shown that the factors affecting

rainfall partitioning patterns include meteorological variables, such as rainfall intensity, air temperature

and potential evapotranspiration (Crockford & Richardson, 2000; Llorens & Domingo, 2007; Staelens

et al., 2008), and the actual contributions are also dependent on vegetation composition and structure

(Crockford & Richardson, 2000; Sadeghi et al., 2020; Zhang et al., 2023). Shrub structures (e.g.,

canopy cover and stand density) strongly affect variations in rainfall partitioning (Chang et al., 2022;

Yue et al., 2021; Zhang et al., 2021). Thinning alters the structures of shrub plots, and can serve as an

important method for regulating the redistribution of water resources in watersheds with shrubland

cover. Thus, examining changes in various components of the shrub water cycle, such as transpiration

and evapotranspiration, as a result of thinning is important to improve understanding of the processes

underlying changes in the water yield.

35

In the present study, changes in rainfall partitioning due to thinning were consistent with a simple

linear relationship between the PAI and canopy water balance values of the two shrub plantations based

on 5 years of measurement data (Figure. 8). Thus, the linear relationship can be used as a useful tool for

predicting net precipitation and observed I rates. When the PAI of the C. korshinskii and S.

psammophila plots was the same, the TF rates of C. korshinskii was significantly higher than that of S.

psammophila, whereas the I rates of C. korshinskii was significantly lower than that of S. psammophila,

there was no significant difference in the SF rates of the C. korshinskii and S. psammophila plots

(Figure. 8). However, in studies conducted in regions with climatic variables similar to those in the

present study, when assessing at the individual plant level, not the stand level, the SF rate of C.

korshinskii was significantly higher than that of S. psammophila and there was no significant difference

in the TF rate (Yuan et al., 2017; Yang et al., 2019b). These findings can be explained by the canopy

projection area of different shrubs. The canopy projection area of C. korshinskii is smaller than that of

S. psammophila. In addition, as the stand density increases, the degree of overestimation of TF and

underestimation of SF and I increases at the individual plant level. As a result, measurements of I at the

individual plant level may underestimate results at the stand level at biome and global scales. Thus, the

role of rainfall partitioning in hydrological processes needs to be studied at the plant community level.

Results from the 3 years after thinning of this study indicate that more heavily thinning treatments had

more pronounced effects on shrub growth, net precipitation, and interception, especially in dry years

(Table 4). However, the effects of thinning intensity are subject to the interplay among various factors.

When shrub plots become dense over time, the canopy intercepts more rain, potentially shortening the

soil water recharge process (Snyder et al., 2021). In water-limited regions, abiotic and biotic factors

affect the soil water content. Vegetation depends on soil moisture, and soil moisture is affected by

36

water uptake by plants, shading, SF, and I (Llorens & Domingo, 2007; Metzger et al., 2017;

Rodriguez-Iturbe, 2000). If a reduction in net precipitation (sum of TF and SF) is not balanced by a

reduction in evaporation due to canopy shading, high densities of shrubs may have a significant impact

on the water budget of the respective ecosystem. Comparative studies of different thinning intensities

can help us to determine the most suitable thinning method for shrub plots to reduce drought stress. For

example, Gebhardt et al. (2014) indicated that repeated moderate thinning was a better option than the

heavy thinning because of heavy thinning induced the progressive development of understory, which

not only competed for resources with trees but also hindered natural regeneration. In this study, the

understory of S. psammophila plot was sparse, so its effects are expected to be minor. Still, for C.

korshinskii plot, the role of the understory in the longer term could become important, affecting the

difference between the two thinning treatments. This further emphasizes the need to examine the

long-term effects of the thinning treatments in the two shrub plots in this study.

4.2 Interception parameters

The S of the C. korshinskii and S. psammophila plots remained around or below 1 mm throughout the

monitoring period (Table 4), which agrees well with ?ndings obtained for other semiarid shrub species

(Zhang et al., 2018). The S of forests is significantly higher than that of shrubs in similar areas. For

example, Ma et al. (2019) reported a S value of 1.34 for Roinia pseudoacacia and 1.43 for Pinus

tabuliformis. Other studies reported that climatic variables (e.g., rainfall intensity and wind speed) and

canopy traits (e.g., cover, height, and leaf area index) influenced S (Carlyle-Moses & Gash, 2011). In

this study, SF was relatively low compared to the values obtained for other parts of rainfall partitioning

in the C. korshinskii and S. psammophila plots (Table 4). The SF-related parameters St and Pt (around

37

0.1 mm) in two shrub plots were much lower than those reported by Zhang et al. (2018) (0.55 mm for

St and 0.68 mm for Pt), but measured SF rates agreed with those found in previous studies in semiarid

regions (Yuan et al., 2017; Yang et al., 2019b; Li et al., 2008; Yue et al., 2021). In this study, the

observed mean values of St and Pt were slightly higher in the C. korshinskii plots than in the S.

psammophila plots, these findings can be explained by the lower threshold of precipitation (0.9 mm for

C. korshinskii vs. 2.1 mm for S. psammophila) and beneficial leaf traits of C. korshinskii versus those

of S. psammophila (Yuan et al., 2017).

In the present study, changes in canopy structure induced by thinning altered interception parameters

(e.g., S, p, and ), thereby influencing I (Table 4). As reported previously, the S and p also lead to E

spatial variability in TF (Loustau et al., 1992; Sun et al., 2015). In this study, in the three thinning years,

there was a greater reduction in S, St, and Pt in the shrub plots subjected to NT and MT than HT.

During 2019?2021, p also increased under NT (0.25 and 0.13 for C. korshinskii and S. psammophila,

respectively) and MT (0.10 and 0.08 for C. korshinskii and S. psammophila, respectively) with

decreasing rainfall. In the plots subjected to HT, p slightly increased and remained at a high level (0.03

for both C. korshinskii and S. psammophila). Both light exposure and aerodynamic conductance of

branches increase with an increase in canopy openness caused by thinning, thus changing

meteorological conditions, such as temperature, humidity, and wind speed, all of which control

evaporation of water intercepted and stored within the canopy (Pypker et al., 2005).

The revised Gash model performance (10% < RE ≤ 20%) improved over the entire observation

period compared to weaker Gash model performance (24% < RE < 84%) for a single year (Table 4).

Fluctuations in model result from the years (see Figure. 9 and Table 6) reflect data availability within a

38

single year rather than vegetation change. Generally, the reliability of derived model parameters

depends on the dataset (i.e., measurement duration) from which the parameters are obtained. In this

study, the parameters derived from three different hydrological years (normal, dry, and extremely dry)

yielded better simulation results than the parameters derived from only one type of hydrological year

(Tables 6 and 7). This clearly shows that the performance of the revised Gash analytical model depends

on reliable parameterization. Including a range of parameters in the revised model from a large number

of hydrological years can enable estimations of rainfall interception over time, which can be used in

shrub water balance studies in arid and semiarid regions.

4.3 Performance of the revised Gash analytical model

The revised Gash analytical model provided good estimates of the total I in both the C. korshinskii and

S. psammophila plots (Tables 6 and 7), and it also captured canopy density variation. Based on the

RMSE, the revised Gash model performed better after thinning than before. In contrast, the NSE values

were markedly better before thinning than after thinning (Table 7). This is not surprising, as intensive

thinning greatly reduces variability in I and TF, and these models are not designed to measure rainfall

events in the open field (Shinohara et al., 2015). In terms of modeling error, the underestimation of

simulated total I using the revised Gash model was similar under the NT treatment in both the C.

korshinskii and S. psammophila plots (Table 6) and within the range reported by other studies (Fan et

al., 2014; Fathizadeh et al., 2018; Junqueira Junior et al., 2019; Limousin et al., 2008; Shinohara et al.,

2015). Muzylo et al. (2009) concluded that the expected modeling error in the prediction of

interception loss can be as high as 20%. In this study, I was slightly overestimated in the control and

thinned plots for small rainfall events during the calibration and validation periods. For high rainfall

39

events, the predicted I was far from the 1:1 line in both the C. korshinskii and S. psammophila plots

(Figure. 10). The revised Gash model severely underestimated I as rainfall increased (Figure. 9). Thus,

this model should be used with caution in areas with high rainfall events (Fathizadeh et al., 2018;

Limousin et al., 2008; Sadeghi et al., 2015). Based on meteorological data collected using automatic

rain gauges in the study area in the past ten years, single rainfall events with heavy rainfall amounts (>

40 mm) account for about 30% of the total rainfall. Therefore, the effect of such rainfall events should

be considered in future research to improve the performance of the model.

The thinning treatments altered the the interception components in the present study. Evaporation

during storm events and the post-storm period accounted for the largest amount of interception loss

before and after thinning (Figure. 11), which is consistent with the findings of previous studies (Sun et

al., 2015; Ma et al., 2019; 2020). Changes in S and strongly affected I (Figure. 12). The degree of E

decrease in the S of the C. korshinskii plots under MT and HT relative to that under NT was smaller

than the degree of decrease in . However, in the S. psammophila plots, the decrease in the S of the E

plots subjected to MT and HT relative to that in the plot subjected to NT was greater than the decrease

in (Table 4). Therefore, S was the main cause of the decrease in I in the thinned plots of C. E

korshinskii, whereas the was the main cause in the thinned plots of S. psammophila. E

Quanti?cation of interception components before and after thinning can help to improve understanding

of changes in interception processes, as well as underlying processes of peak ?ows, in?ltration, and

water resources in this ecosystem.

Typically, model error refers to the results of model validation rather than calibration (Muzylo et al.,

2009). This paper shows performance of the revised Gash model in S. psammophila plots than in C.

40

korshinskii plots during the validation period. This indicates the revised Gash model for interception

loss modeling can be more appropriate for the S. psammophila plots than for the C. korshinskii plots in

terms of RE and NSE values (Table 7). As noted in previous studies, the revised version has been

extensively applied in semiarid climates where sparse vegetation cover is common, and performs well

(Fan et al., 2014; Muzylo et al., 2009). Interception modeling is important for estimating the water

balance and thus plays a crucial role in hydrological simulations and determining eco-hydrological

services (Junqueira Junior et al., 2019). Future work should focus on operationalizing canopy

interception models for routine use by shrub managers to optimize shrub management strategies.

5. Conclusions

Thinning increased net precipitation reaching the soil surface and reduced canopy interception. In the

plots subjected to HT, signi?cant effects of the extremely dry year could not be identi?ed in rainfall

partitioning measurements and canopy parameters, such as the PAI and S. This suggests that HT allows

both C. korshinskii and S. psammophila plots to adapt better to dry weather conditions.

The interception parameters (S, c, Pt, St, and ) decreased with thinning intensity, and changes in S E

and strongly affected simulated I. Of note, the decrease in simulated I in the thinned plots of C. E

korshinskii was mainly caused by the S, whereas it was mainly caused by the in the S. E

psammophila plots.

The revised analytical Gash model provided a good estimate of the total I of the two shrub species (RE

< 20%). However, when evaluating the I of individual rainfall events, the simulation ability and

accuracy of the model decreased significantly, with moderate overestimation of low rainfall events (<

41

10 mm) and severe underestimation of high rainfall events (> 40 mm). These over- and

underestimations were mainly caused by uncertainty in the input parameters. Using data from a type

hydrological year or growing season to derive canopy parameters is not recommended. Parameters that

can reliably estimate I can only be generated using data with varying rainfall amounts. Moreover, as

intensive thinning greatly reduces the variability in I, the revised analytical Gash model performed

better after thinning than before thinning based on the RE values.

Although the effects of thinning on rainfall partitioning in this study lasted for three years, the effective

length of time for thinning remains to be addressed. In addition, further studies should also evaluate the

effects of other important water cycle elements (soil moisture, transpiration, etc.) to address the effects

on the whole water cycle and to improve the implementation of tree plantations in water-limited

regions.

Conflicts of interests

The authors declare that there are no conflicts of interest.

Acknowledgements

This work was supported by the Strategic Priority Research Program of the Chinese Academy of

Sciences (XDA23070202) and the National Natural Science Foundation of China (No. 41977016).

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